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yes, but the implementation depends on what kind of backend software you are using. The general method for doing this involves scripting your 404 page so that it parses out the extra path information for the stuff not found (ie, /Sample/One_Two.html) and then either performs a lookup or, in your case, a string manipulation (ie, lower(url)) and then redirects to the new URL.
But, without knowing what kind of server side scripting you have going on, its hard to say exactly. At any rate, that is the general technique. It can also be used to provide search engine friendly URL's when you have dynamic sites. PM me if you have more specific questions.
rob
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