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Pagerank, anchor text, and relevance to your niche, to name a few. It's hard to say exactly because no one knows all the factors, and they're interrelated. For instance, if you have a site about dogs and you get a link from a PR5 page about turtles, it may not help you as much as a PR3 link from a page about dogs. In other words, don't obsess over high PR links that have no relevance to your site.
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